Senior Paleontologist, Curator

(701) 328-8163
600 East Boulevard
Bismarck, ND 58505-0840

Job Description

Clint manages the paleontology program for the North Dakota Geological Survey and serves as curator of the North Dakota State Fossil Collection. His research activities focus on Eocene and Oligocene vertebrate faunas and the anatomy and systematic relationships of ornithischian dinosaurs.


2012: Ph.D. Geosciences, The University of Texas at Austin
2005: B.S. Geology & Geophysics / Zoology (Double Major), University of Wisconsin-Madison


Wang, X., Emry, R.J., Boyd, C.A., Person, J.J., White, S.C., and Tedford, R.H. 2023. An exquisitely preserved skeleton of Eoarctos vorax (nov. gen. et sp.) from Fitterer Ranch, North Dakota (Early Oligocene) and systematics and phylogeny of North American early arctoids (Carnivora, Caniformia). Society of Vertebrate Paleontology Memoir 22. Journal of Vertebrate Paleontology. 42(2, Supplement). DOI: 10.1080/02724634.2022.2145900

Boyd, C.A., 2023.  A Significant Fossil Collection Comes Home.  Geo News 50(1):6-9.

Korth, W.W., Boyd, C.A., Emry, R.J., and Person, J.J. 2023. Additional small mammals from the Oligocene Brule Formation (late Orellan-Whitneyan) of southwestern North Dakota. Paludicola. 14(2):57-74.

Barnes, B., and Boyd, C.A., 2022.  Small Exhibits, Big Impact.  Geo News 49(2):16-18.

Drumheller, S.K., Boyd, C.A., Barnes, B.M.S., and Householder, M.L. 2022. Biostratinomic alterations of an Edmontosaurus “mummy” reveal a pathway for soft tissue preservation without invoking “exceptional conditions.” PLoS ONE. 17(10):e0275240. DOI: 10.1371/journal.pone.o275240

Salcido, C.J., Wilson, P., Tweet, J.S., McCann, B.E., Boyd, C.A., and Santucci, V.L. 2022.Theodore Roosevelt National Park: Paleontological resource inventory (public version). Natural Resource Report NPS/THRO/NRR—2022/2385. National Park Service, Fort Collins, Colorado. DOI: 10.36967/nrr-2293509

Korth, W.W., Boyd, C.A., Person, J.J., and Anderson, D.K. 2022. Fossil mammals from ant mounds situated on exposures of the Big Cottonwood Creek Member of the Chadron Formation (latest Eocene-early Oligocene, Sioux county, Nebraska. Paludicola. 13:191-344.

Boyd, C.A., 2021.  A Heavy Burden to Find a Tiny Lizard.  Geo News 48(2):13-15.

Madzia, D., Arbour, V.M., Boyd, C.A., Farke, A.A., Cruzado-Caballero, P., and Evans, D.C. 2021. The phylogenetic nomenclature of ornithischian dinosaurs. PeerJ. 9:e12362. DOI: 10.7717/peerj.12362

VanVranken, N., and Boyd, C.A. 2021. The first in situ collection of a mosasaurine from the marine Breien Member of the Hell Creek Formation in south-central North Dakota, USA. PaleoBios. 28:1-11.

Korth, W.W., Boyd, C.A., Emry, R.J., and Person, J.J. 2021. Marsupials (Mammalia, Metatheria) from the Brule Formation (Oligocene) North Dakota. Journal of Palaeontology. 95(1):193-204. DOI: 10.1017/jpa.2020.41

Boyd, C.A., 2021.  Pioneering Fossil Finds from North Dakota.  Geo News 48(1):14-17.

Welsh, E., Boyd, C.A., and Spearing, K. 2020. Split carinae on a specimen of false saber-toothed cat (Carnivora: Nimravidae) and the implications for analogous tooth abnormality formation in mammals and theropod dinosaurs. Proceedings of the South Dakota Academy of Science. 99:69-82.

Czaplewski, N.J., Person, J.J., Boyd, C.A., and Emry, R.J. 2019. A new species of bat (Chiroptera: Vespertillionidae) form the early Oligocene global cooling period, Brule Formation, North Dakota, USA. Palaeo Vertebrata. 42:e2. DOI: 10.18563/pv.42.2.e2

Korth, W.W., Boyd, C.A., and Person, J.J. 2019. Whitneyan (middle Oligocene) rodents from Obritsch Ranch (Stark County, North Dakota) and a review of Whitneyan rodent fossil record. Annals of Carnegie Museum. 85:249-278. DOI: 10.2992/007.085.0304

Korth, W.W., Emry, R.J., Boyd, C.A., and Person, J.J. 2019. Rodents (Mammalia) from Fitterer Ranch, Brule Formation (Oligocene), North Dakota. Smithsonian Contributions to Paleontology. 103:1-45.

Boyd, C.A., 2019.  Public Fossil Digs Program Surges to Record Level.  Geo News 46(1):10-12.

Welsh, E., Pagnac, D., and Boyd, C.A. 2018. New material of Amynodontidae (Perrissodactyla) from Badlands National Park and its implications on the Whitneyan-Arikareean transition. Proceedings of the South Dakota Academy of Science. 97:181-197.

Lawver, D.R., and Boyd, C.A. 2018. An avian eggshell from the Brule Formation (Oligocene) of North Dakota. Journal of Vertebrate Paleontology. 38:e1486848. DOI: 10.1080/02724634.2018.1486848

Boyd, C.A., 2018.  A Pleasing Discovery from North Dakota's Ancient Seas.  Geo News 45(2):5-10.

Boyd, C.A., and Webster, J. R. 2018. Depositional History of the Chadron Formation in North Dakota. North Dakota Geological Survey Report of Investigation 120:1-114.

Madzia, D., Boyd, C.A., and Mazuch, M. 2018. A basal ornithopod dinosaur from the Cenomanian of the Czech Republic. Journal of Systematic Palaeontology. 16(11):967-979 DOI: 10.1080/14772019.2017.1371258

Boyd, C.A., Person, J.J., Barnes, B., 2017.  Additions to the Lancian Mammalian Fauna from southwest North Dakota.  Journal of Vertebrate Paleontology DOI: 10.1080/02724634.2017.1325368

Boyd, C.A., 2017.  Paleontological Tourism in North Dakota.  Geo News 44(2):20-23.

Boyd, C.A., 2017. A New Addition To The Cretaceous Seaway Of North Dakota. Geo News 44(1):20-23.

Boyd, C.A., and J.J. Person. 2016. New Additions To The Lancian Mammalian Fauna From Southwest North Dakota. Journal of Vertebrate Paleontology (2016 Program and Abstracts).

Boyd, C.A., J.J. Person, and B. Barnes. 2016. New Additions To The Lancian Mammalian Fauna From Southwest North Dakota, Poster.

Person, J.J., and C.A. Boyd. 2016. New Paleontology Exhibits Opened Across North Dakota. Geo News 43(2):1-5.

Boyd, C.A., 2016. The Overlooked Marvelous Reptile. Geo News 43(1):2-6. 3:e1263

Boyd, C.A. 2015. Insight on the anatomy, systematic relationships, and age of the Early Cretaceous ankylopollexian dinosaur Dakotadon lakotaensis. PeerJ. 3:e1263

Boyd, C.A. 2015. In pursuit of early mammals by Zofia Kielan-Jaworowska. Quarterly Review of Biology. 90:207-208.

Clarke, J.A., and Boyd, C.A. 2015. Methods for the Quantitative Comparison of Molecular Estimates of Clade Age and the Fossil Record. Systematic Biology. 64(1):25-41.

Boyd, C.A. 2014. The cranial anatomy of the neornithischian dinosaur Thescelosaurus neglectus. PeerJ. 2:e669

Boyd, C.A., Weiler, M.W., Householder, M.L., and Schumaker, K.K. 2014. The diversity of cimolestan mammals within the White River Group of South Dakota and Nebraska. Acta Palaeontologica Polonica. 59(4):771-778.

Boyd, C.A., and Welsh, E. 2014. Description of the first Orellan fauna defined in Badlands National Park, Interior, South Dakota and implications for the stratigraphic position of the Bloom Basin Limestone Bed. Dakoterra. 6:124-147.

Boyd, C.A., Drumheller, S.K., and Gates, T.A. 2013. Crocodyliform feeding traces on juvenile ornithischian dinosaurs from the Upper Cretaceous (Campanian) Kaiparowits Formation, Utah. PLOS ONE. 8(2):e57605

Gates, T.A., Lund, E.K., Boyd, C.A., Getty, M.A., DeBlieux, D.D., Kirkland, J.I., and Evans, D.C. 2013. Ornithopod dinosaurs from the Grand Staircase-Escalante National Monument Region, Utah, and their role in paleobiogeographic and macroevolutionary studies. In: Titus, A., and Loewen, M.A. (Eds.), At the Top of the Grand Staircase: The Late Cretaceous in Utah. Indiana University Press. p. 463-481.

Ksepka, D.T., and Boyd, C.A. 2012. A new method for quantifying historical trends in the completeness of the fossil record and the contributing factors: an example using Aves. Paleobiology. 38(1):112-125.

Brown, C.M., Boyd, C.A., and Russell, A.P. 2011. A new basal neornithischian from the Frenchman Formation, Saskatchewan, Canada and its associated fauna: implications for Late Maastrichtian ornithischian diversity in North America. Zoological Journal of the Linnean Society. 163(4):1157-1198.

Boyd, C.A., Cleland, T.P., and Novas, F. 2011. Osteogenesis, homology, and function of intercostal plates on ornithischian dinosaurs. Zoomorphology. 130(4):305-313.

Boyd, C.A., Cleland, T. P., Marrero, N. L., and Clarke, J. A. 2011. Exploring the effects of phylogenetic uncertainty and consensus trees on stratigraphic consistency scores: a new program and a standardized method proposed. Cladistics. 27(1):52-60.

Boyd, C.A., Brown, C.M., Scheetz, R., and Clarke, J.A. 2009. Taxonomic revision of the basal neornithischian taxa Thescelosaurus and Bugenasaura. Journal of Vertebrate Paleontology. 29(3):758-770.

Clarke, J.A., Ksepka, D.T., Stucchi, M., Urbina, M., Giannini, N., Bertelli, S., Narváez, Y., and Boyd, C.A. 2007. Paleogene equatorial penguins challenge the proposed relationship between penguin biogeography, body size evolution, and Cenozoic climate change. Proceedings of the National Academy of Sciences. 104(28):11545-11550.